Under Construction

Unplaced Acalyptratae

Keith Bayless
Containing group: Schizophora


Three genera of flies in the Schizophora, Mallochianamyia Santos-Neto, Paraleucopis Malloch, and Schizostomyia Malloch, are unplaced to family. Wheeler (2000) suggested that these genera may form a monophyletic clade. This assemblage is sometimes referred to as the "Schizostomyia group" (Hamilton et al. 2006). Wheeler (2000) and Hamilton et al. (2006) mention that there is an undescribed genus of flies from Australia related to these three genera.

Schizostomyia and Mallochianamyia are from Chile (Wheeler 2000, Thompson 2005) and Paraleucopis is found in Mexico and the Southwestern United States. Paraleucopis larvae are inhabitants of bird's nests (McAlpine 1987).

Discussion of Phylogenetic Relationships

Paraleucopis has been associated with Chamaemyiidae, but McAlpine (1987) admits that this placement was provisional. Though Paraleucopis shares some characteristics with Chamaemyiidae, its genitalic configuration is strongly divergent from the rest of the family. Morphological data suggest that this group may be related to Asteiidae, particularly the genus Leiomyza (McAlpine 1989).


Hamilton, J., D. Yeates, A. Hastings, D. Colless, D. McAlpine, D. Bickel, G. Daniels, M. Schneider, P. Cranston, and S. Marshall. 2006. On the fly: the interactive atlas and key to Australia fly families. (CD-ROM) Australian Biological Resources Study (ABRS) and Centre for Biological Information Technology (CBIT).

McAlpine, J.F. 1987. 88. Chamaemyiidae. Manual Nearct. Dipt. 2:965-971.

McAlpine, J.F. 1989. 116. Phylogeny and classification of the Muscomorpha. Manual Nearct. Dipt. 3:1397-1518.

Wheeler T. A. 2000. Systematics of Mallochianamyia Santos-Neto (Diptera: Schizophora), Neotropical acalyptrate flies of uncertain family relationships. Insect Syst. Evol. 31:209-224.

About This Page

Keith Bayless
North Carolina State University, Raleigh, North Carolina, USA

Correspondence regarding this page should be directed to Keith Bayless at

All Rights Reserved.

Citing this page:

Bayless, Keith. 2008. Unplaced Acalyptratae. Version 25 March 2008 (under construction). http://tolweb.org/Unplaced_Acalyptratae/119717/2008.03.25 in The Tree of Life Web Project, http://tolweb.org/

edit this page
close box

This page is a Tree of Life Branch Page.

Each ToL branch page provides a synopsis of the characteristics of a group of organisms representing a branch of the Tree of Life. The major distinction between a branch and a leaf of the Tree of Life is that each branch can be further subdivided into descendent branches, that is, subgroups representing distinct genetic lineages.

For a more detailed explanation of the different ToL page types, have a look at the Structure of the Tree of Life page.

close box

Unplaced Acalyptratae

Page Content

articles & notes



Explore Other Groups

random page

  go to the Tree of Life home page